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The story of the original manga begins when a girl named Emi uses a pencil that can grant any wish. Emi gets caught up in the robbery of a convenience store, and her life is in danger. She uses the pencil to wish for a "hero that will save everyone." Four heroes appear in response to Emi's wish to change the course of the future. The anime will have a different story from the original manga.

This is a story about the strongest single dog in history searching for his future wife. He is acknowledged as the strongest existence and stands at the top of the world, but wealth, fame, and power are not worth mentioning in his eyes. The strongest apostle has escaped and appeared on the human continent, and the reason is shockingly, Don’t misunderstand! I’m here to find my future wife. this is a story about the strongest single dog searching for a wife in history.

In the year 2043, , the world's first successful full-dive VRMMO was released. In addition to its ability to perfectly simulate the five senses, along with its many other amazing features, the game promised to offer players a world full of infinite possibilities. Nearly two years later, soon-to-be college freshman, Reiji Mukudori, is finally able to buy a copy of the game and start playing. With some help from his experienced older brother, Shuu, and his partner Embryo, Reiji embarks on an adventure into the world of . Just what will he discover and encounter in this game world known for its incredible realism and infinite possibilities?

In the year 2043, , the world's first successful full-dive VRMMO was released. In addition to its ability to perfectly simulate the five senses, along with its many other amazing features, the game promised to offer players a world full of infinite possibilities. Nearly two years later, soon-to-be college freshman, Reiji Mukudori, is finally able to buy a copy of the game and start playing. With some help from his experienced older brother, Shuu, and his partner Embryo, Reiji embarks on an adventure into the world of . Just what will he discover and encounter in this game world known for its incredible realism and infinite possibilities?

The story revolves around the IS Academy and its students who are training to become pilots for the IS (Infinite Stratos) weapon system. It so happens that this system can only be operated by females. However there was one young man named Orimura Ichika (15) who was able to use the IS. He then finds himself attending the busy academy dominated by girls. As it turns out Shinonono Houki, a childhood friend, is one of his new classmates.

Second season of Infinite Stratos.

Ichika has gone home for the holidays. On one particular day, his rest is cut short when the girls start showing up at his doorstep one after the other.

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A project was launched where scientists discovered a mysterious substance that can interface directly with the human brain. This substance, called "Exotic Matter (XM)," has existed since ancient times, influencing human minds and the progress of humanity. In the wake of this discovery, a battle of powerful nations and corporate giants has been unleashed. Organizations across the globe have embarked on a secret race to exploit XM. It represents both an opportunity and a threat to humanity. Two Factions seeks to control the XM. The Enlightened view XM's power as a gift that enhances human experience and discovery. The Resistance sees XM as a hostile takeover of the human mind, choosing technology as humanity's best path forward. XM, and the mystery behind it lie at the center of this battle for the fate of humanity. 2018—Now, a new struggle is about to unfold in Tokyo and across the globe. Dangerous and powerful forces seeking to exploit the potential of XM will collide. This groundbreaking project will mark the beginning of an epic augmented-reality experience combining the three elements of animation, location-based gaming, and the real world.

A project was launched where scientists discovered a mysterious substance that can interface directly with the human brain. This substance, called "Exotic Matter (XM)," has existed since ancient times, influencing human minds and the progress of humanity. In the wake of this discovery, a battle of powerful nations and corporate giants has been unleashed. Organizations across the globe have embarked on a secret race to exploit XM. It represents both an opportunity and a threat to humanity. Two Factions seeks to control the XM. The Enlightened view XM's power as a gift that enhances human experience and discovery. The Resistance sees XM as a hostile takeover of the human mind, choosing technology as humanity's best path forward. XM, and the mystery behind it lie at the center of this battle for the fate of humanity. 2018—Now, a new struggle is about to unfold in Tokyo and across the globe. Dangerous and powerful forces seeking to exploit the potential of XM will collide. This groundbreaking project will mark the beginning of an epic augmented-reality experience combining the three elements of animation, location-based gaming, and the real world.

This is simply the OVA summing up the major events and battles of "Initial D 4th Stage" much like the "Initial D Battle Stage" original summed up the first 3 seasons. Additionally, has 2 extra battles from the manga that were not incorporated into the original 4th Stage.