Akito doesn't want to fight. Despite a childhood spent on the anime Gekiganger 3, a Mecha show, he'd rather cook than pilot a Mecha. Fate intervenes when his home on Mars is destroyed, and he is transported instantly to the Earth, mysteriously. He has questions no one can answer fully, but follows a girl from a chance meeting in hopes to discover any. The girl, Yurika, is captain of the private battleship Nadesico, and in order to follow her, he enlists as their cook. Possessing the nanite implants that allow to control mechas, he's a handy backup pilot for the mechas of the Nadesico. He joins a crew bent on avenging Mars that seems to be composed of only misfits, otakus, and ditzes; however, in reality, they are handpicked experts. They take their own private war back to Mars to face the harsh reality that life may not always be like a Giant Mecha series.

Akito doesn't want to fight. Despite a childhood spent on the anime Gekiganger 3, a Mecha show, he'd rather cook than pilot a Mecha. Fate intervenes when his home on Mars is destroyed, and he is transported instantly to the Earth, mysteriously. He has questions no one can answer fully, but follows a girl from a chance meeting in hopes to discover any. The girl, Yurika, is captain of the private battleship Nadesico, and in order to follow her, he enlists as their cook. Possessing the nanite implants that allow to control mechas, he's a handy backup pilot for the mechas of the Nadesico. He joins a crew bent on revenging Mars and composed of misfits, otakus, and ditzes. But in reality they are handpicked experts. They take their own private war back to Mars to face the harsh reality that life may not always be like a Giant Mecha Series.

Two years have passed since the end of the "Martian Successor Nadesico." Akito and Yurika have disappeared. The Jovians and the Earthlings have joined forces and Ruri is now captain of Nadesico B. As the popularity of Boson Jumping grows, a vast transportation network has been developed. Dubbed the Hisago Plan, this network of Chulip portals holds the answers to the mysteries behind Bose particles and their power. If those secrets should fall into the wrong hands, it could mean big trouble. And big trouble there is. A Martian splinter group has launched an offensive, leaving the United Forces befuddled and desperate. So much so, in fact, that the former crew of the Nadesico is called back to active duty for a special unsanctioned mission-to save us all. Does this menagerie of misfits have what it takes? Who will win the race for the Boson technology? And what the heck happened to Akito and Yurika?

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